生物專業(yè)英語第三版蔣悟生編__課文翻譯

1、生物工程專業(yè)英語oneInside the Living Cell: Structure andFunction of Internal Cell Parts1、 Cytoplasm: The Dynamic, Mobile Factory細胞質(zhì)：動力工廠Most of the properties we associate with life are properties of the cytoplasm. Much of the mass of a cell consists of this semifluid substance, which is bounded on the outs

2、ide by the plasma membrane. Organelles are suspended within it, supported by the filamentous network of the cytoskeleton. Dissolved in the cytoplasmic fluid are nutrients, ions, soluble proteins, and other materials needed for cell functioning.生命的大部分特征表現(xiàn)在細胞質(zhì)的特征上。細胞質(zhì)大部分由半流體物質(zhì)組成，并由細胞膜（原生質(zhì)膜）包被。細胞器懸浮在其中

3、，并由絲狀的細胞骨架支撐。細胞質(zhì)中溶解了大量的營養(yǎng)物質(zhì)，離子，可溶蛋白以及維持細胞生理需求的其它物質(zhì)。2、The Nucleus: Information Central（細胞核：信息中心）The eukaryotic cell nucleus is the largest organelle and houses the genetic material (DNA) on chromosomes. (In prokaryotes the hereditary material is found in the nucleoid.) The nucleus also contains one o

4、r two organelles-the nucleoli-that play a role in cell division. A pore-perforated sac called the nuclear envelope separates the nucleus and its contents from the cytoplasm. Small molecules can pass through the nuclear envelope, but larger molecules such as mRNA and ribosomes must enter and exit via

5、 the pores.真核細胞的細胞核是最大的細胞器，細胞核對染色體組有保護作用（原核細胞的遺傳物質(zhì)存在于擬核中）。細胞核含有一或二個核仁，核仁促進細胞分裂。核膜貫穿許多小孔，小分子可以自由通過核膜，而象mRNA和核糖體等大分子必須通過核孔運輸。3、Organelles: Specialized Work Units（細胞器：特殊的功能單位）All eukaryotic cells contain most of the various kinds of organelles, and each organelle performs a specialized function in the

6、cell. Organelles described in this section include ribosomes, the endoplasmic reticulum, the Golgi complex, vacuoles, lysosomes, mitochondria, and the plastids of plant cells.所有的真核細胞都含有多種細胞器，每個細胞器都有其特定功能。本節(jié)主要介紹核糖體，內(nèi)質(zhì)網(wǎng)，高爾基體系，液泡，溶酶體，線粒體和植物細胞中的質(zhì)體。The number of ribosomes within a cell may range from a f

7、ew hundred to many thousands. This quantity reflects the fact that, ribosomes are the sites at which amino acids are assembled into proteins for export or for use in cell processes. A complete ribosome is composed of one larger and one smaller subunit. During protein synthesis the two subunits move

8、along a strand of mRNA, "reading" the genetic sequence coded in it and translating that sequence into protein. Several ribosomes may become attached to a single mRNA strand; such a combination is called a polysome. Most cellular proteins are manufactured on ribosomes in the cytoplasm. Expo

9、rtable proteins and membrane proteins are usually made in association with the endoplasmic reticulum.核糖體的數(shù)量變化從幾百到幾千，核糖體是氨基酸組裝成蛋白質(zhì)的重要場所。完整的核糖體由大亞基和小亞基組成。核糖體沿著mRNA移動并閱讀遺傳密碼，翻譯成蛋白質(zhì)。一條mRNA上可能有多個核糖體，稱多聚核糖體。大多數(shù)細胞蛋白是由細胞質(zhì)中核糖體生產(chǎn)。輸出蛋白和膜蛋白通常與內(nèi)質(zhì)網(wǎng)有關(guān)。The endoplasmic reticulum, a lacy array of membranous sacs, tub

10、ules, and vesicles, may be either rough (RER) or smooth (SER). Both types play roles in the synthesis and transport of proteins. The RER, which is studded with polysomes, also seems to be the source of the nuclear envelope after a cell divides.內(nèi)質(zhì)網(wǎng)，帶有花邊的生物囊，有管狀，泡狀之分，以及光滑和粗糙面區(qū)別。兩種都與蛋白質(zhì)的合成和運輸有關(guān)。粗糙內(nèi)質(zhì)網(wǎng)上分

11、布許多核糖體，也可能提供細胞分裂后所需的細胞膜。SER lacks polysomes; it is active in the synthesis of fats and steroids and in the oxidation of toxic substances in the cell. Both types of endoplasmic reticulum serve as compartments within the cell where specific products can be isolated and subsequently shunted to particul

12、ar areas in or outside the cell.光滑內(nèi)質(zhì)網(wǎng)上無核糖體，主要作用是脂肪和類固醇的合成以及細胞內(nèi)有毒物質(zhì)的氧化。兩種內(nèi)質(zhì)網(wǎng)合成的產(chǎn)物在其中進行分流或運輸?shù)郊毎狻ransport vesicles may carry exportable molecules from the endoplasmic reticulum to another membranous organelle, the Golgi complex. Within the Golgi complex molecules are modified and packaged for export

13、out of the cell or for delivery else where in the cytoplasm.運輸小泡能夠?qū)⒖蛇\輸分子從內(nèi)質(zhì)網(wǎng)運輸?shù)礁郀柣鶑?fù)合體上。在高爾基復(fù)合體中修飾，包裝后輸出細胞或傳遞到細胞質(zhì)中的其他場所。Vacuoles in cells appear to be hollow sacs but are actually filled with fluid and soluble molecules. The most prominent vacuoles appear in plant cells and serve as water reservoirs

14、and storage sites for sugars and other molecules. Vacuoles in animal cells carry out phagocytosis (the intake of particulate matter) and pinocytosis (vacuolar drinking).細胞中的液泡好象是中空的，但實際上充滿了液體和可溶分子。最典型的液泡存在于植物細胞中，儲備水，糖以及其它分子。動物中的液泡起吞噬和胞飲作用。 A subset of vacuoles are the organelles known as lysosomes,

15、which contain digestive enzymes (packaged in lysosomes in the Golgi complex) that can break down most biological macromolecules. They act to digest food particles and to degrade damaged cell parts.溶酶體是液泡亞單位，含有消化酶，降解大部分生物大分子。消化食物微粒和降解損傷的細胞殘片。Mitochondria are the sites of energy-yielding chemical reac

16、tions in all cells. In addition, plant cells contain plastids that utilize light energy to manufacture carbohydrates in the process of photosynthesis. It is on the large surface area provided by the inner cristae of mitochondria that ATP-generating enzymes are located. Mitochondria are self-replicat

17、ing, and probably they are the evolutionary descendants of what were once free-living prokaryotes.線粒體是細胞中化學(xué)產(chǎn)能的場所。另外，植物細胞中的質(zhì)體在光合作用中利用光能產(chǎn)生碳水化合物，線粒體內(nèi)嵴上提供了很大的表面積并分布著產(chǎn)ATP酶。線粒體自我復(fù)制，并且可能是自由生活的原核生物在進化中形成的后代。There are two types of plastids: leucoplasts, which lack pigments and serve as storage sites for star

18、ch, proteins, and oils; and chromoplasts, which contain pigments. The most important chromoplasts are chloroplasts-organelles that contain the chlorophyll used in photosynthesis. The internal structure of chloroplasts includes stacks of membranes called grana, which are embedded in a matrix called t

19、he stroma.質(zhì)體有兩種類型：白色體，缺乏色素，是淀粉，蛋白質(zhì)和油的儲備場所；色質(zhì)體，含有色素。葉綠體是最重要的色質(zhì)體，含有與光合作用有關(guān)的葉綠素。葉綠體的內(nèi)部結(jié)構(gòu)是由多層膜形成的葉綠體基粒，其中包埋在基質(zhì)中的基粒稱子座。4、The Cytoskeleton（細胞骨架）All eukaryotic cells have a cytoskeleton, which is a convoluted latticework of filaments and tubules that appears to fill all available space in the cell and prov

20、ides support for various other organelles. A large portion of the cytoskeleton consists of threadlike microfilaments composed mainly of the contractile protein actin. They are involved in many types of intracellular movements in plant and animal cells. A second protein, myosin, is involve in the con

21、traction of muscle cells. Another main structural component of the cytoskeleton consists of microtubules, which are composed of the globular protein tubulin and together act as scaffolding that provides a stable cell shape. Cytoskeletal intermediate filaments appear to impart tensile strength to the

22、 cell cytoplasm. Mechanoenzymes such as myosin, dynein, and kinesin interact with the cytoskeletal filaments and tubules to generate forces that cause movements.所有的細胞都有細胞骨架，網(wǎng)絡(luò)結(jié)構(gòu)的纖絲充滿了它所能觸及的全部空間并且對細胞器提供支持作用。細胞骨架大部分由微絲組成，微絲主要由可收縮的肌動蛋白組成。動植物細胞的許多種類型細胞內(nèi)運動與肌動蛋白有關(guān)。第二類蛋白是肌球蛋白，它與肌肉細胞的收縮有關(guān)。細胞骨架的另一個主要結(jié)構(gòu)成分是微管，

23、由球狀的微管蛋白組成，象腳手架一般維持細胞的穩(wěn)定形態(tài)。細胞骨架的中間絲提供了細胞質(zhì)伸縮動力。機械酶，例如，肌球蛋白，動力蛋白，驅(qū)動蛋白與微絲，微管相互作用產(chǎn)生動力而引起細胞運動。5、Cellular Movements（細胞運動 Although the cytoskeleton provides some stability穩(wěn)固 to cells, its microtubules英maikrutju:bju:l and filaments 絲狀物 and their associated proteins enable cells to move by creeping 爬行 or gli

24、ding 滑動. Such movements require a solid固體的 substrate to which the cell can adhere附著 and can be guided by the geometry dimitri  幾何形狀of the surface. Some cells also exhibit izibit 展覽 chemotaxis, kemtæksis趨藥性 the ability to move toward or away from the source of a diffusing 擴散 chemical.盡管細胞骨架

25、提供了細胞的某些穩(wěn)定性，微絲，微管及相關(guān)蛋白能使細胞爬行或滑動。這種運動需要固體基質(zhì)依托并通過表面幾何形狀的改變而運動。某些細胞具備趨藥性，即趨向或逃離擴散開的化學(xué)源。Certain eukaryotic cells can swim freely in liquid environments, propelled by whiplike cilia sili纖毛or flagella 英fldel鞭毛. Both cilia and flagella have the same internal structure: nine doublets (pairs of microtubules)

26、 are arranged in a ring and extend the length of the cilium or flagellum, and two more microtubules run down the center of the ring. Every cilium or flagellum grows only from the cell surface where a basal body基體is located. Movement is based on the activities of tiny dynein動力蛋白 side arms that extend

27、 from one of the microtubules of each doublet成對的東西.某些真核細胞能在液體液體中自由運動，由纖毛或鞭毛推動。纖毛和鞭毛具有同樣的內(nèi)部結(jié)構(gòu)：九個雙微管環(huán)形排列，縱向延伸，環(huán)中心是兩個或以上微管組成。纖毛或鞭毛從細胞表面的基體出生長，雙微管的動力蛋白臂從一側(cè)延伸到另一側(cè)而引起運動。Nutrients, proteins, and other materials within most plant cells are moved about via cytoplasmic streaming. The process occurs as myosin

28、maiusin肌凝蛋白 proteins attached to organelles 細胞器,小器官 push against microfilaments arrayed rei 展示throughout the cell. Microfilaments and microtubules are responsible for almost all major cytoplasmic movements. During cell division, microtubules of the spindle assembled 集合, 收集 from tubutin subunits near

29、 organelles called centrioles move the chromosomes. 大部分植物細胞的營養(yǎng)，蛋白質(zhì)和其它物質(zhì)由細胞質(zhì)流運輸。這個過程是由于依附在細胞器上的肌球蛋白反推排列在細胞周圍的微絲形成的。絕大部分細胞質(zhì)運動由微絲和微管完成。在細胞分裂期間，中心粒周圍的由微管蛋白亞基裝配形成的紡錘體微管移向染色體。課后作業(yè)：第一第二篇閱讀材料答案：1B,2C,3B,4B,5D,6c; 1A,2B,3D,4A38twoPhotosynthesisPhotosynthesis occurs only in the chlorophyllchlorophyll葉綠素-conta

30、ining cells of green plants, algae藻, and certain protists 原生生物and bacteria. Overall, it is a process that converts light energy into chemical energy that is stored in the molecular bonds. From the point of view of chemistry and energetics, it is the opposite of cellular respiration. Whereas 然而 cellu

31、lar細胞的 respiration 呼吸is highly exergonic吸收能量的 and releases energy, photosynthesis光合作用 requires energy and is highly endergonic.光合作用只發(fā)生在含有葉綠素的綠色植物細胞，海藻，某些原生動物和細菌之中?？傮w來說，這是一個將光能轉(zhuǎn)化成化學(xué)能，并將能量貯存在分子鍵中，從化學(xué)和動能學(xué)角度來看，它是細胞呼吸作用的對立面。細胞呼吸作用是高度放能的，光合作用是需要能量并高吸能的過程。Photosynthesis starts with CO2 and H2O as raw mater

32、ials and proceeds through two sets of partial reactions. In the first set, called the light-dependent reactions, water molecules are split裂開 (oxidized), 02 is released, and ATP and NADPH are formed. These reactions must take place in the presence of 在面前 light energy. In the second set, called light-

33、independent reactions, CO2 is reduced (via the addition of H atoms) to carbohydrate. These chemical events rely on the electron carrier NADPH and ATP generated by the first set of reactions.光合作用以二氧化碳和水為原材料并經(jīng)歷兩步化學(xué)反應(yīng)。第一步，稱光反應(yīng)，水分子分解，氧分子釋放，ATP和NADPH形成。此反應(yīng)需要光能的存在。第二步，稱暗反應(yīng)，二氧化碳被還原成碳水化合物，這步反應(yīng)依賴電子載體NADPH以及第

34、一步反應(yīng)產(chǎn)生的ATP。Both sets of reactions take place in chloroplasts. Most of the enzymes and pigments 色素for the lightdependent reactions are embedded 深入的 內(nèi)含的in the thylakoid類囊體 membrane膜 隔膜 of chloroplasts 葉綠體. The dark reactions take place in the stroma.基質(zhì) 兩步反應(yīng)都發(fā)生在葉綠體中。光反應(yīng)需要的大部分酶和色素包埋在葉綠體的類囊體膜上。暗反應(yīng)發(fā)生在基質(zhì)中。

35、1、How Light Energy Reaches Photosynthetic Cells（光合細胞如何吸收光能的）The energy in light photons in the visible part of the spectrum can be captured by biological molecules to do constructive work. The pigment chlorophyll in plant cells absorbs photons within a particular absorption spectrums statement of th

36、e amount of light absorbed by chlorophyll at different wavelengths. When light is absorbed it alters the arrangement of electrons in the absorbing molecule. The added energy of the photon boosts the energy condition of the molecule from a stable state to a less-stable excited state. During the light

37、-dependent reactions of photosynthesis, as the absorbing molecule returns to the ground state, the "excess" excitation energy is transmitted to other molecules and stored as chemical energy.生物分子能捕獲可見光譜中的光能。植物細胞中葉綠素在不同光波下吸收部分吸收光譜。在吸收分子中，光的作用使分子中的電子發(fā)生重排。光子的能量激活了分子的能量狀態(tài)，使其從穩(wěn)定態(tài)進入不穩(wěn)定的激活態(tài)。All ph

38、otosynthetic organisms contain various classes of chlorophylls and one or more carotenoid (accessory) pigments that also contribute to photosynthesis. Groups of pigment molecules called antenna complexes are present on thylakoids. Light striking any one of the pigment molecules is funneled to a spec

39、ial chlorophyll a molecule, termed a reaction-center chlorophyll, which directly participates in photosynthesis. Most photosynthetic organisms possess two types of reaction-center chlorophylls, P680 and P700, each associated with an electron acceptor molecule and an electron donor. These aggregation

40、s are known respectively as photosystem (P700) and photosystem (P680).所有的光合作用生物含有不同等級的葉綠素和一個或多個類胡蘿卜素（光合作用的輔助色素）。稱作天線復(fù)合體的色素分子群存在于類囊體中。激活色素分子的光能進入葉綠素反應(yīng)中心，其直接參與光合作用。大部分光反應(yīng)細胞器擁有兩套反應(yīng)中心，P680和P700，每個光系統(tǒng)都含有一個電子受體和電子供體。這些集合體就是大家熟識的光合系統(tǒng)和光合系統(tǒng) 。2、The Light-Dependent Reaction: Converting Solar Energy into Chemic

41、al-Bond Energy光反應(yīng)：光能轉(zhuǎn)化成化學(xué)鍵能The photosystems of the light-dependent reactions are responsible for the packaging of light energy in the chemical compounds ATP and NADPH. This packaging takes place through a series of oxidation reduction reactions set in motion when light strikes the P680 reaction cent

42、er in photosystem . In this initial event water molecules are cleaved, oxygen is released, and electrons are donated. These electrons are accepted first by plastoquinone and then by a series of carriers as they descend an electron transport chain. For each four electrons that pass down the chain, tw

43、o ATPs are formed. The last acceptor in the chain is the P700 reaction center of photosystem . At this point incoming photons boost the energy of the electrons, and they are accepted by ferredoxin. Ferredoxin is then reoxidized, and the coenzyme NADP+ is reduced to the NADPH. The ATP generated previ

44、ously and the NADPH then take part in the light independent reactions.光反應(yīng)的光系統(tǒng)將光能轉(zhuǎn)化成化學(xué)復(fù)合物ATP和NADPH。當(dāng)光激活光系統(tǒng)的光反應(yīng)中心時，通過一系列的氧化還原反應(yīng)實現(xiàn)能量的傳遞。反應(yīng)開始時，水被分解，氧被釋放并提供電子。電子首先傳遞給質(zhì)體醌，然后通過一系列載體形成的電子傳遞鏈。每傳遞4個電子，形成2個ATP。最后一個受體存在于光反應(yīng)系統(tǒng)的反應(yīng)中心里。此處光子激活電子，電子傳遞給鐵氧還蛋白。鐵氧還蛋白再氧化，并且輔酶NADP+還原成NADPH。早期產(chǎn)生的ATP和NADPH進入暗反應(yīng)。The productio

45、n of ATP from the transport of electrons excited by light energy down an electron transport chain is termed photophosphorylation. The one-way flow of electrons through photosystems II and I is called noncyclic photophosphorylation; plants also derive additional ATP through cyclic photophosphorylatio

46、n, in which some electrons are shunted back through the electron transport chain between photosystems and .由電子傳遞鏈偶連產(chǎn)生ATP的過程稱為光合磷酸化。通過光合系統(tǒng)流經(jīng)光合系統(tǒng)的電子路徑稱非循環(huán)式光合磷酸化；植物通過循環(huán)式光合磷酸化獲得額外的ATP，一些電子在光合系統(tǒng)和之間的電子傳遞鏈中回流。3、The Light-Independent Reactions: Building Carbohydrates暗反應(yīng)：碳水化合物的形成In the light-independent reac

47、tions of photosynthesis, which are driven by ATP and NADPH, C02 is converted to carbohydrate. The reactions are also known as the Calvin-Benson cycle. Atmospheric CO2, is fixed as it reacts with ribulose biphosphate (RuBP), a reaction that is catalyzed by the enzyme ribulose biphosphate carboxylase.

48、 The reduction Of C02 to carbohydrate (fructose diphosphate) is completed via several more steps of the cycle. Finally, RUBP is regenerated so that the cycle may continue.由ATP和NADPH驅(qū)動的暗反應(yīng)中，二氧化碳轉(zhuǎn)化成碳水化合物。即卡爾文循環(huán)。二磷酸核酮糖固定二氧化碳，由二磷酸核酮糖羧化酶催化。4、Oxygen: An Inhibitor of photosynthesis（氧：光合作用的抑制因子）High levels

49、of oxygen in plant cells can disrupt photosynthesis and can also cause photorespiration-an inefficient fun of the dark reactions in which 02 is fixed rather than C02 and no carbohydrate is produced.Reprieve from Photorespiration: The C4 PathwayMost plants are C3 plants; they experience decreased car

50、bohydrate production under hot, dry conditions as a result of the effects of photorespiration. Among C4 plants, however, special leaf anatomy and a unique biochemical pathway enable the plant to thrive in and conditions. Thus C4 plants lessen photorespiration by carrying out photosynthesis only in c

51、ells that are insulated from high levels of CO2. They also possess a novel mechanism for carbon fixation. 大部分植物是碳3植物，在高溫干旱條件下，由于光呼吸作用而使碳水化合物的合成降低。而在大多數(shù)的碳4植物中，由于葉脈的特殊構(gòu)造和獨特的化學(xué)路徑使植物依然很茂盛。這是碳固定的一個新機制。課后作業(yè)：第一篇閱讀材料答案：1B,2A,3A,4D,5C,6cthreeCellular Reproduction: Mitosis and Meiosis1、The Nucleus and Chromos

52、omesThe cell nucleus is the main repository of genetic information. Within the nucleus are the chromosomes tightly coiled strands of DNA and clusters of associated proteins. Long stretches of the continuous DNA molecule wind around these clusters of proteins, or histones, forming beadlike complexes

53、known as nucleosomes. More coiling and supercoiling produces a dense chromosome structure. Each long strand of DNA combines with histones and nonhistone proteins to make up the substance chromatin.細胞核是貯藏遺傳信息的主要場所。DNA盤繞成螺旋線以及相關(guān)的成簇蛋白質(zhì)。DNA螺旋線纏繞成簇的組蛋白形成珠鏈狀的核小體。這些螺旋和超螺旋形成致密的染色體組結(jié)構(gòu)。每個長鏈DNA與組蛋白和非組蛋白一起構(gòu)成染色質(zhì)

54、物質(zhì)。A pictorial display of an organism's chromosomes in the coiled, condensed state is known as a karyotype. Karyotype reveal that in most cells all but sex chromosomes are present as two copies, referred to as homologous pairs. Non-sex chromosomes are called autosomes. Organisms whose cells cont

55、ain two sets of parental chromosomes are called diploid; those with cells containing a single set of parental chromosomes are called haploid. 染色體致密的超螺旋狀態(tài)我們稱染色體組。除了性染色體外，大多數(shù)細胞的染色體組成對出現(xiàn)，稱同源染色體對。非性染色體稱常染色體。生物細胞含有兩套父母本染色體的稱二倍體；含有單套染色體的稱單倍體。2、The Cell CycleThe cell cycle is a regular sequence in which th

56、e cell grows, prepares for division, and divides to form two daughter cells, each of which then repeats the cycle. Such cycling in effect makes single-celled organisms immortal. Many cells in multicellular organisms, including animal muscle and nerve cells, either slow the cycle or break out of it a

57、ltogether.在細胞生長過程中，細胞循環(huán)遵循特定程序，分裂準(zhǔn)備，分裂成2個子細胞，子細胞再循環(huán)。此循環(huán)使得單細胞永生。多細胞生物中的許多細胞，包括動物肌肉和神經(jīng)細胞，要么降低循環(huán)速度，要么同時分裂。The normal cell cycle consists of four phases. The first three include G1, the period of normal metabolism; S phase, during which normal synthesis of biological molecules continues, DNA is replicate

58、d, and histones are synthesized; and G2, a brief period of metabolism and additional growth. Together the G1, S, and G2 phases are called interphase. The fourth phase of the cell cycle is M phase, the period of mitosis, during which the replicated chromosomes condense and move and the cell divides.

59、It is believed that properties of the cell cytoplasm control the cell cycle, along with external stimulators and inhibitors such as chalones.正常細胞循環(huán)由4個時期組成。頭三期包括G1,正常新陳代謝；S期,正常新陳代謝同時，DNA復(fù)制，組蛋白合成； G2 期，短期的新陳代謝和少許生長。G1, S, 和G2稱分裂間期。最后是M期，有絲分裂期，復(fù)制的染色體組濃縮，移動并細胞分裂。據(jù)稱是染色質(zhì)控制了細胞循環(huán)，伴隨外部激活因子和抑制因子如抑素。 3、Mitosis: